Statistics and Breed-wide Allele Frequency - Biewer

Updated Oct 2, 2026

DLA Class I Haplotype Frequencies

DLA1 #STR typesBiewer (n=142)Biewer Yorshire Terrier (n=54)Biewer Terrier (n=236)Yorkshire Terrier (n=16)Biro Biewer (n=3)
1005389 371 277 1810.0250.0090.013----
1007380 372 281 1820.0040.009--0.03--
1008386 373 289 1820.007----0.06--
1012388 369 289 1880.2180.1570.2290.280.2
1014375 373 287 1780.0280.0090.0570.09--
1016382 371 277 1780.0210.0190.0110.03--
1019380 373 287 1850.0740.0830.0610.09--
1030380 373 293 1780.4650.3520.4920.310.5
1035386 373 277 184--0.009------
1040380 371 277 1860.0920.2040.078----
1043393 381 277 1830.0070.0090.011----
1046376 379 291 1800.028--0.021----
1068380 373 287 181--0.009------
1087380 371 277 1780.004--------
1092376 379 277 181--0.037--0.06--
1104386 373 289 1860.0110.0190.017----
1105382 379 277 1780.004------0.3
1115386 371 277 182----0.002----
1117376 373 277 180----0.002----
1118376 377 277 1800.0040.019------
1130380 373 287 1780.0070.009------
1131388 379 277 1810.0040.0460.0060.03--

DLA Class II Haplotype Frequencies

DLA2 #STR typesBiewer (n=142)Biewer Yorshire Terrier (n=54)Biewer Terrier (n=236)Yorkshire Terrier (n=16)Biro Biewer (n=3)
2003343 324 2820.2540.2130.2480.380.2
2005339 322 2800.0390.0090.021----
2013345 327 2840.0460.1390.064----
2016339 323 2840.0810.0930.0720.09--
2021339 324 2680.004--------
2023341 323 2820.4650.3520.4940.310.5
2037341 327 2800.0280.0090.0570.09--
2040345 327 2800.0600.0830.0250.03--
2044343 324 268--0.037--0.06--
2053343 324 280--0.009------
2074341 324 2840.0040.0190.002----
2080339 325 2760.0040.009--0.03--
2081343 322 2820.0110.0190.017----
2082339 325 268--0.009------
2083339 324 2820.004------0.3
2084339 323 2680.004--------

Allele Frequencies

#Locus NameAlleleBiewer (n=142)Biewer Yorshire Terrier (n=54)Biewer Terrier (n=236)Yorkshire Terrier (n=25)Biro Biewer (n=3)
1AHT121 92----0.0150.040.2
940.004--0.0110.06--
980.1090.1480.1040.20--
1000.2040.0650.1190.20--
1020.2990.3060.3520.120.7
1040.2150.3610.2200.08--
1060.032--0.0130.08--
1080.1060.0830.1230.20--
1100.0070.0280.0190.02--
1120.0250.0090.021----
122----0.002--0.2
124----0.002----
2AHT137 1310.2360.2780.2440.200.3
133------0.08--
1370.3270.2780.3090.40--
1390.1270.1200.110----
143--0.009------
147----0.0110.04--
1490.004--0.004----
1510.2430.2690.2560.20--
1530.0630.0460.0660.060.7
155------0.02--
3AHTH130 1170.011--0.011----
1190.3660.3890.3350.140.7
1210.1650.1570.0910.080.2
1230.0210.0740.0170.08--
125----0.0020.02--
1270.3380.2690.4240.240.2
1290.0920.1020.1040.40--
1310.0070.0090.0170.02--
133------0.02--
4AHTh171-A 217----0.002----
2190.6480.7780.7140.500.3
2250.1970.0930.1310.120.5
227----0.0020.04--
2290.004--0.0020.04--
2310.1090.1110.1140.120.2
2330.0280.0190.0280.14--
2350.011--0.006----
2370.004----0.04--
5AHTh260 2380.0460.0740.0210.22--
240------0.02--
2420.0420.0190.0250.12--
2440.0850.0370.0720.020.2
2460.2570.2410.2750.360.3
2480.0740.0650.112----
2500.4960.5370.4890.260.5
254--0.0280.004----
6AHTk211 870.4720.5460.4870.210.7
890.018--0.0210.04--
910.4120.4350.3710.650.3
950.0210.0090.0570.06--
970.0770.0090.0640.04--
7AHTk253 2840.0490.0090.0550.020.3
2860.0460.0280.0570.02--
2880.1940.3430.2200.42--
2900.1480.2690.1290.24--
2920.5530.3520.5380.260.7
2940.011----0.04--
8C22.279 1160.1130.1110.0890.020.2
1180.0740.0370.0360.12--
1200.0040.0090.0080.04--
1220.0110.0090.0080.04--
1240.1830.1940.1930.04--
1260.3350.4170.4340.460.8
1280.2640.2220.2200.18--
1300.018--0.0110.10--
9FH2001 124------0.12--
1320.4790.4720.5250.400.2
136--0.009--0.02--
140----0.0080.12--
1440.0850.0930.0510.080.2
1480.4230.3980.3980.240.7
1520.0140.0090.0170.02--
156--0.019------
10FH2054 1480.2220.2590.2060.34--
1520.1270.0560.1590.260.5
1560.3490.4440.3540.320.5
1600.2040.1390.197----
1640.0600.0460.0610.02--
1680.0390.0560.0230.06--
11FH2848 2320.011--0.002----
2340.0920.0560.076----
2360.3200.3330.3370.24--
2380.1730.1300.1590.220.5
2400.3870.3980.3830.400.5
242----0.002----
2440.0180.0830.0400.14--
12INRA21 910.1900.1940.212--0.2
950.2010.1480.1990.420.3
970.2890.2590.1690.220.3
99--0.028--0.10--
1010.3100.3430.4050.260.2
1050.0110.0280.015----
13INU005 1060.0110.0560.0470.06--
1100.063--0.0400.10--
120----0.002----
1240.8170.7960.7900.561.0
1260.0490.0930.0320.08--
128----0.0130.02--
1300.018--0.0190.12--
1320.0420.0560.0570.04--
134------0.02--
14INU030 1440.2180.2500.2100.280.3
1460.0350.0280.034----
1480.0560.0830.0400.10--
1500.5320.5650.6530.420.5
1520.0810.0280.0110.080.2
1540.0770.0460.0530.10--
156------0.02--
15INU055 208------0.04--
2100.5250.5090.6120.460.3
2120.0280.0460.0280.06--
2140.1800.1300.1120.20--
2160.0670.0930.0700.02--
2180.2010.2220.1780.220.7
16LEI004 850.0810.1020.1120.18--
950.8240.8060.7460.700.8
970.0880.0930.1400.02--
101------0.02--
1050.004------0.2
1070.004--0.0020.08--
17REN105L03 2290.011--------
2310.0210.0830.0340.08--
2330.3240.2310.3050.08--
2350.1230.1300.1460.08--
2370.2990.2310.2690.24--
2410.2220.3240.2460.501.0
247------0.02--
18REN162C04 1980.004----0.14--
2000.0250.0460.0570.06--
2020.0950.1020.0890.120.3
2040.0490.0740.0570.040.3
2060.3200.2780.2420.060.3
2080.0350.0190.0530.06--
2100.4720.4810.5020.52--
19REN169D01 2020.2080.2040.1590.160.2
210--0.0190.0130.08--
2120.3420.3700.4220.180.2
2160.2680.2870.2780.420.7
2200.1830.1200.1290.16--
20REN169O18 158------0.04--
1600.042--0.0300.16--
1620.1690.2220.1380.40--
1640.6440.4810.6840.301.0
1660.0920.1850.102----
1680.0320.0280.0380.02--
1700.0210.0830.0080.06--
172------0.02--
21REN247M23 2680.2750.2410.1970.24--
2700.0210.0560.0360.22--
2720.3910.4260.4220.320.2
276--------0.3
2780.3130.2780.3450.220.5
22REN54P11 2220.0140.1020.0300.04--
2240.007--0.006----
2260.2180.2130.2310.060.2
230------0.02--
2320.2430.1760.1670.340.2
2340.0950.0370.0850.020.3
2360.2610.3430.3390.500.3
2380.1620.1300.1420.02--
23REN64E19 1390.1730.2410.1400.220.7
1430.014--0.008----
1450.4080.2690.4790.06--
1470.3700.4630.3090.56--
1490.0180.0190.0320.040.3
1510.007--0.0060.02--
1530.007--0.0250.08--
1550.0040.009--0.02--
24VGL0760 120.2460.4440.2780.140.3
130.0110.0370.011----
140.0390.0280.040----
20.20.0280.0190.0400.040.3
21.20.3350.2500.3200.36--
22.20.0320.0280.0720.14--
23.20.1510.0740.1270.24--
24.20.1550.1200.1100.080.3
25.20.004--0.002----
25VGL0910 15.1------0.02--
16.10.0630.0280.1210.16--
17.10.004--0.0040.08--
18.10.0210.0090.0360.04--
190.0810.0650.030----
19.10.2150.3060.2690.100.3
200.0250.056--0.06--
20.10.0110.0090.0210.160.2
210.004--------
21.10.1870.1300.2010.080.3
220.0320.0190.049----
22.10.3490.3610.2610.100.2
23----0.002----
23.10.007--0.0060.14--
24.10.0040.019--0.06--
26VGL1063 8----0.0020.02--
9------0.02--
11------0.02--
120.021--0.021----
130.021--0.0150.04--
140.2540.1480.2730.360.2
150.3980.4540.4090.320.8
160.1270.1670.1210.02--
170.0460.0650.0490.10--
180.0920.1390.0660.10--
190.0070.0090.002----
200.0350.0190.038----
21----0.004----
27VGL1165 15------0.02--
18------0.02--
190.4820.4540.5610.320.8
200.2430.1570.1120.14--
210.0770.0650.0550.060.2
220.004--------
250.0040.019--0.08--
260.0250.0740.0320.06--
27----0.004----
280.0850.0740.0830.26--
290.0770.1480.1330.04--
300.0040.0090.015----
31----0.004----
28VGL1828 140.1270.1110.083----
150.0420.0830.025--0.2
160.2010.2410.1630.280.3
170.1160.1480.1440.120.3
18--0.0090.0210.040.2
190.3660.2500.3580.26--
200.0460.0190.0190.02--
210.046--0.0510.14--
220.0560.1390.1330.10--
23----0.0020.04--
29VGL2009 100.011--0.0080.04--
110.2430.1940.2750.12--
120.028--0.0020.020.2
130.5700.6480.5210.440.3
140.0250.0460.0780.260.5
150.1130.1110.1000.12--
160.011--0.015----
30VGL2409 130.0140.0190.0040.020.2
140.007--0.0170.02--
150.007--0.0060.06--
160.0920.0830.1550.08--
170.2320.3150.2250.400.2
180.3100.2310.3390.380.7
190.011--0.0130.02--
200.3270.3520.2350.02--
21----0.006----
31VGL2918 120.0770.1760.1190.140.2
130.1200.0090.1480.140.2
140.0740.1110.0760.12--
150.025--0.0300.10--
160.0040.0090.015----
17.30.5250.4630.4680.280.7
18------0.02--
18.30.0850.1110.0320.14--
19.30.0920.1020.1000.06--
20.3--0.019------
22.3----0.013----
32VGL3008 130.0530.0560.034----
14----0.002----
150.1300.0930.1570.140.3
160.0350.1020.0640.02--
170.1230.2040.0720.080.2
180.4190.2780.3450.320.2
190.0740.0280.1020.16--
200.1650.2410.2120.240.3
21----0.002----
22----0.0110.04--
33VGL3235 120.0040.0090.0130.04--
130.1200.0650.1060.04--
140.2360.2780.1950.380.2
150.1060.0930.0950.06--
160.0420.0370.0420.12--
170.2570.1390.2290.100.3
180.1550.1850.1840.160.5
190.0040.0090.0210.04--
200.0280.0090.0110.02--
210.0490.1760.1040.04--

Standard genetic assessment based on 33 autosomal STR loci

Biewer

  NNaNeHoHeF
Mean4598.3943.5200.6730.6850.017
 SE 0.4200.1850.0200.0190.005

Biewer Yorshire Terrier

  NNaNeHoHeF
Mean546.0003.3880.6900.672-0.029
 SE 0.2870.1730.0230.0210.014

Biewer Terrier

  NNaNeHoHeF
Mean2367.3033.4110.6590.6730.021
 SE 0.3720.1900.0210.0200.008

Yorkshire Terrier

  NNaNeHoHeF
Mean246.8483.9060.6930.7200.038
 SE 0.2910.2260.0180.0140.019

Biro Biewer

  NNaNeHoHeF
Mean32.6362.2110.5860.483-0.190
 SE 0.1470.1360.0570.0350.076

Standard genetic assessment based on 7 STRs in the DLA region

Biewer

  NNaNeHoHeF
Mean4596.7142.5700.5980.597-0.001
 SE 0.8010.1710.0310.0300.005

Biewer Yorshire Terrier

  NNaNeHoHeF
Mean546.1433.0840.6960.660-0.060
 SE 0.6520.2440.0260.0300.027

Biewer Terrier

  NNaNeHoHeF
Mean2365.2862.3820.5720.567-0.006
 SE 0.6910.1510.0350.0310.010

Yorkshire Terrier

  NNaNeHoHeF
Mean245.5712.8940.5420.6360.159
 SE 0.4450.2320.0560.0330.066

Biro Biewer

  NNaNeHoHeF
Mean32.4292.0960.7620.460-0.535
 SE 0.2750.2310.1460.0830.116

Standard genetic assessment for individual STR loci

Biewer

#LocusNNaNeHoHeF
1AHT121459124.9600.8150.798-0.021
2AHT137459104.2290.7870.764-0.031
3AHTH13045993.5540.7040.7190.020
4AHTh171-A45991.9680.4890.4920.006
5AHTh26045983.0850.6570.6760.029
6AHTk21145952.5230.5990.6040.007
7AHTk25345962.9300.6520.6590.010
8C22.27945983.8110.7260.7380.016
9FH200145982.4400.5780.5900.020
10FH205445964.1940.7630.762-0.002
11FH284845973.4460.7200.710-0.014
12INRA2145963.8930.7130.7430.040
13INU00545991.5880.3540.3700.043
14INU03045972.4510.6020.592-0.017
15INU05545962.6290.6000.6200.032
16LEI00445961.6030.3760.3760.000
17REN105L0345974.0620.7520.7540.002
18REN162C0445973.0980.6500.6770.040
19REN169D0145953.6140.7200.7230.005
20REN169O1845982.2790.5480.5610.024
21REN247M2345953.1110.6890.679-0.016
22REN54P1145984.5730.7130.7810.087
23REN64E1945983.0940.6130.6770.094
24VGL076045994.5820.7260.7820.071
25VGL0910459155.1710.8200.807-0.016
26VGL1063459133.8600.7170.7410.032
27VGL1165459133.1760.6670.6850.026
28VGL1828459105.1030.8040.804-0.000
29VGL200945972.6700.6110.6250.023
30VGL240945994.0090.7520.751-0.002
31VGL2918459113.6330.6910.7250.046
32VGL3008459104.7060.7890.788-0.002
33VGL3235459106.1840.8200.8380.022

Biewer Yorshire Terrier

#LocusNNaNeHoHeF
1AHT1215473.8800.7960.742-0.073
2AHT1375464.1130.7590.757-0.003
3AHTH1305463.7870.7590.736-0.032
4AHTh171-A5441.5970.3700.3740.009
5AHTh2605472.7890.6110.6410.047
6AHTk2115442.0490.5000.5120.023
7AHTk2535453.1830.6480.6860.055
8C22.2795473.6400.7040.7250.030
9FH20015462.5600.5740.6090.058
10FH20545463.4210.6670.7080.058
11FH28485453.3730.7780.704-0.106
12INRA215464.0670.7040.7540.067
13INU0055441.5410.3890.351-0.107
14INU0305462.5500.5930.6080.025
15INU0555452.9740.6480.6640.024
16LEI0045431.4970.3520.332-0.059
17REN105L035454.2380.6670.7640.127
18REN162C045463.0550.7410.673-0.101
19REN169D015453.6250.8520.724-0.176
20REN169O185453.0940.7040.677-0.040
21REN247M235443.1290.7960.680-0.170
22REN54P115464.5000.7410.7780.048
23REN64E195452.9000.7590.655-0.159
24VGL07605483.5300.7960.717-0.111
25VGL091054104.0080.7780.751-0.036
26VGL10635473.5780.7590.721-0.054
27VGL11655483.7290.7220.7320.013
28VGL18285485.5120.8150.8190.005
29VGL20095442.1170.5930.528-0.123
30VGL24095453.5240.6850.7160.043
31VGL29185483.5600.8330.719-0.159
32VGL30085475.0150.8330.801-0.041
33VGL323554105.6790.8520.824-0.034

Biewer Terrier

#LocusNNaNeHoHeF
1AHT121236124.6840.7920.786-0.007
2AHT13723674.2130.8010.763-0.050
3AHTH13023683.2120.6570.6890.046
4AHTh171-A23681.8490.4700.459-0.025
5AHTh26023672.9910.6480.6660.026
6AHTk21123652.6130.6190.617-0.002
7AHTk25323652.7690.6440.639-0.008
8C22.27923683.5230.6990.7160.024
9FH200123652.2850.5250.5620.066
10FH205423664.2410.7710.764-0.009
11FH284823673.4100.6950.7070.017
12INRA2123653.6070.7120.7230.015
13INU00523681.5800.3430.3670.065
14INU03023662.1030.5300.525-0.010
15INU05523652.3540.5720.5750.005
16LEI00423641.7000.4150.412-0.009
17REN105L0323654.0260.7580.752-0.009
18REN162C0423663.0510.6140.6720.086
19REN169D0123653.3680.6690.7030.048
20REN169O1823662.0000.4960.5000.008
21REN247M2323642.9660.6270.6630.054
22REN54P1123674.4540.7250.7750.066
23REN64E1923672.8880.5590.6540.144
24VGL076023694.6250.6990.7840.108
25VGL0910236114.9880.8140.800-0.018
26VGL1063236113.7690.7290.7350.008
27VGL116523692.8030.6480.643-0.008
28VGL1828236104.8960.8090.796-0.017
29VGL200923672.7480.6310.6360.008
30VGL240923694.0800.7970.755-0.055
31VGL291823693.6590.6690.7270.079
32VGL3008236104.7720.7540.7900.046
33VGL3235236106.3320.8430.842-0.001

Yorkshire Terrier

#LocusNNaNeHoHeF
1AHT1212496.5450.8800.847-0.039
2AHT1372473.9680.8000.748-0.070
3AHTH1302483.9810.6800.7490.092
4AHTh171-A2473.2980.8000.697-0.148
5AHTh2602463.8340.6800.7390.080
6AHTk2112452.1370.5420.532-0.018
7AHTk2532463.2890.8000.696-0.149
8C22.2792483.6550.6800.7260.064
9FH20012473.9430.6400.7460.143
10FH20542453.4530.7600.710-0.070
11FH28482443.5010.6000.7140.160
12INRA212443.3070.7200.698-0.032
13INU0052482.8540.6400.6500.015
14INU0302463.5510.6400.7180.109
15INU0552463.2720.7600.694-0.094
16LEI0042451.8880.3600.4700.235
17REN105L032463.0560.7200.673-0.070
18REN162C042473.1570.7200.683-0.054
19REN169D012453.7540.6800.7340.073
20REN169O182473.5510.7600.718-0.058
21REN247M232443.8940.7200.7430.031
22REN54P112472.6880.4800.6280.236
23REN64E192472.6710.7200.626-0.151
24VGL07602464.2660.6400.7660.164
25VGL091024118.8650.7600.8870.143
26VGL10632493.9180.7600.745-0.020
27VGL11652494.8640.6800.7940.144
28VGL18282485.1650.8000.8060.008
29VGL20092463.4250.6000.7080.153
30VGL24092483.1650.6000.6840.123
31VGL29182486.0390.7200.8340.137
32VGL30082474.6820.8400.786-0.068
33VGL323524104.8830.7600.7950.044

Biro Biewer

#LocusNNaNeHoHeF
1AHT121332.0000.6670.500-0.333
2AHT137321.8000.6670.444-0.500
3AHTH130332.0000.3330.5000.333
4AHTh171-A332.5710.3330.6110.455
5AHTh260332.5711.0000.611-0.636
6AHTk211321.8000.6670.444-0.500
7AHTk253321.8000.6670.444-0.500
8C22.279321.3850.3330.278-0.200
9FH2001332.0000.6670.500-0.333
10FH2054322.0001.0000.500-1.000
11FH2848322.0001.0000.500-1.000
12INRA21343.6001.0000.722-0.385
13INU005311.0000.0000.0000.000
14INU030332.5710.6670.611-0.091
15INU055321.8000.0000.4441.000
16LEI004321.3850.3330.278-0.200
17REN105L03311.0000.0000.0000.000
18REN162C04333.0000.6670.6670.000
19REN169D01332.0000.6670.500-0.333
20REN169O18311.0000.0000.0000.000
21REN247M23332.5711.0000.611-0.636
22REN54P11343.6000.6670.7220.077
23REN64E19321.8000.0000.4441.000
24VGL0760333.0000.6670.6670.000
25VGL0910343.6001.0000.722-0.385
26VGL1063321.3850.3330.278-0.200
27VGL1165321.3850.3330.278-0.200
28VGL1828343.6000.6670.7220.077
29VGL2009332.5710.6670.611-0.091
30VGL2409332.0000.6670.500-0.333
31VGL2918332.0000.6670.500-0.333
32VGL3008343.6001.0000.722-0.385
33VGL3235332.5711.0000.611-0.636

Standard genetic assessment for 7 STRs in the DLA region

Biewer

#LocusNNaNeHoHeF
1DLA I-3CCA45982.2230.5420.5500.014
2DLA I-4ACA45992.3880.5900.581-0.016
3DLA I-4BCT45963.2470.6950.692-0.004
4DLA1131459102.9330.6540.6590.008
55ACA45942.8110.6580.644-0.021
65ACT45952.6130.6080.6170.015
75BCA45951.7710.4360.435-0.001

Biewer Yorshire Terrier

#LocusNNaNeHoHeF
1DLA I-3CCA5482.0390.5930.510-0.163
2DLA I-4ACA5473.0610.7410.673-0.100
3DLA I-4BCT5453.4370.6670.7090.060
4DLA11315494.1100.7410.7570.021
55ACA5443.5070.7960.715-0.114
65ACT5453.0360.7220.671-0.077
75BCA5452.3960.6110.583-0.049

Biewer Terrier

#LocusNNaNeHoHeF
1DLA I-3CCA23682.1810.5250.5420.030
2DLA I-4ACA23652.1740.5510.540-0.020
3DLA I-4BCT23653.0150.6950.668-0.040
4DLA113123682.6380.6310.621-0.017
55ACA23642.5480.6100.607-0.004
65ACT23642.4670.6100.595-0.026
75BCA23631.6530.3810.3950.034

Yorkshire Terrier

#LocusNNaNeHoHeF
1DLA I-3CCA2463.6000.6250.7220.135
2DLA I-4ACA2482.8940.6250.6550.045
3DLA I-4BCT2453.4290.6250.7080.118
4DLA11312463.5450.6670.7180.071
55ACA2442.5890.6250.614-0.018
65ACT2452.3090.2500.5670.559
75BCA2451.8920.3750.4710.204

Biro Biewer

#LocusNNaNeHoHeF
1DLA I-3CCA332.5711.0000.611-0.636
2DLA I-4ACA332.5711.0000.611-0.636
3DLA I-4BCT332.5711.0000.611-0.636
4DLA1131321.3850.3330.278-0.200
55ACA332.5711.0000.611-0.636
65ACT322.0001.0000.500-1.000
75BCA311.0000.0000.0000.000