Statistics and Breed-wide Allele Frequency - Biewer

Updated Sep 2, 2025

DLA Class I Haplotype Frequencies

DLA1 #STR typesBiewer (n=139)Biewer Yorshire Terrier (n=54)Biewer Terrier (n=231)Yorkshire Terrier (n=16)Biro Biewer (n=3)
1005389 371 277 1810.0250.0090.013----
1007380 372 281 1820.0040.009--0.03--
1008386 373 289 1820.007----0.06--
1012388 369 289 1880.2230.1570.2270.280.2
1014375 373 287 1780.0290.0090.0580.09--
1016382 371 277 1780.0220.0190.0110.03--
1019380 373 287 1850.0720.0830.0610.09--
1030380 373 293 1780.4570.3520.4960.310.5
1035386 373 277 184--0.009------
1040380 371 277 1860.0940.2040.078----
1043393 381 277 1830.0070.0090.011----
1046376 379 291 1800.029--0.022----
1068380 373 287 181--0.009------
1087380 371 277 1780.004--------
1092376 379 277 181--0.037--0.06--
1104386 373 289 1860.0110.0190.013----
1105382 379 277 1780.004------0.3
1115386 371 277 182----0.002----
1117376 373 277 180----0.002----
1118376 377 277 1800.0040.019------
1130380 373 287 1780.0070.009------
1131388 379 277 1810.0040.0460.0060.03--

DLA Class II Haplotype Frequencies

DLA2 #STR typesBiewer (n=139)Biewer Yorshire Terrier (n=54)Biewer Terrier (n=231)Yorkshire Terrier (n=16)Biro Biewer (n=3)
2003343 324 2820.2590.2130.2470.380.2
2005339 322 2800.0400.0090.022----
2013345 327 2840.0470.1390.065----
2016339 323 2840.0790.0930.0710.09--
2021339 324 2680.004--------
2023341 323 2820.4570.3520.4980.310.5
2037341 327 2800.0290.0090.0580.09--
2040345 327 2800.0610.0830.0240.03--
2044343 324 268--0.037--0.06--
2053343 324 280--0.009------
2074341 324 2840.0040.0190.002----
2080339 325 2760.0040.009--0.03--
2081343 322 2820.0110.0190.013----
2082339 325 268--0.009------
2083339 324 2820.004------0.3
2084339 323 2680.004--------

Allele Frequencies

#Locus NameAlleleBiewer (n=139)Biewer Yorshire Terrier (n=54)Biewer Terrier (n=231)Yorkshire Terrier (n=25)Biro Biewer (n=3)
1AHT121 92----0.0150.040.2
940.004--0.0110.06--
980.1010.1480.1060.20--
1000.2090.0650.1150.20--
1020.3020.3060.3510.120.7
1040.2120.3610.2210.08--
1060.032--0.0130.08--
1080.1080.0830.1260.20--
1100.0070.0280.0190.02--
1120.0250.0090.019----
122----0.002--0.2
124----0.002----
2AHT137 1310.2300.2780.2400.200.3
133------0.08--
1370.3270.2780.3140.40--
1390.1290.1200.108----
143--0.009------
147----0.0110.04--
1490.004--0.004----
1510.2480.2690.2580.20--
1530.0610.0460.0650.060.7
155------0.02--
3AHTH130 1170.011--0.011----
1190.3670.3890.3380.140.7
1210.1690.1570.0910.080.2
1230.0220.0740.0170.08--
125----0.0020.02--
1270.3350.2690.4200.240.2
1290.0900.1020.1040.40--
1310.0070.0090.0170.02--
133------0.02--
4AHTh171-A 217----0.002----
2190.6400.7780.7100.500.3
2250.2010.0930.1320.120.5
227----0.0020.04--
2290.004--0.0020.04--
2310.1120.1110.1170.120.2
2330.0290.0190.0280.14--
2350.011--0.006----
2370.004----0.04--
5AHTh260 2380.0470.0740.0220.22--
240------0.02--
2420.0430.0190.0260.12--
2440.0860.0370.0710.020.2
2460.2630.2410.2710.360.3
2480.0760.0650.113----
2500.4860.5370.4940.260.5
254--0.0280.004----
6AHTk211 870.4710.5460.4870.210.7
890.018--0.0220.04--
910.4210.4350.3700.650.3
950.0180.0090.0580.06--
970.0720.0090.0630.04--
7AHTk253 2840.0500.0090.0560.020.3
2860.0470.0280.0560.02--
2880.1980.3430.2190.42--
2900.1510.2690.1300.24--
2920.5430.3520.5390.260.7
2940.011----0.04--
8C22.279 1160.1150.1110.0910.020.2
1180.0760.0370.0370.12--
1200.0040.0090.0090.04--
1220.0110.0090.0090.04--
1240.1870.1940.1970.04--
1260.3200.4170.4260.460.8
1280.2700.2220.2210.18--
1300.018--0.0110.10--
9FH2001 124------0.12--
1320.4860.4720.5240.400.2
136--0.009--0.02--
140----0.0090.12--
1440.0790.0930.0520.080.2
1480.4210.3980.3980.240.7
1520.0140.0090.0170.02--
156--0.019------
10FH2054 1480.2270.2590.2080.34--
1520.1220.0560.1560.260.5
1560.3530.4440.3550.320.5
1600.1980.1390.195----
1640.0610.0460.0630.02--
1680.0400.0560.0240.06--
11FH2848 2320.011--0.002----
2340.0860.0560.078----
2360.3200.3330.3380.24--
2380.1760.1300.1560.220.5
2400.3880.3980.3850.400.5
242----0.002----
2440.0180.0830.0390.14--
12INRA21 910.1940.1940.212--0.2
950.2010.1480.2010.420.3
970.2950.2590.1690.220.3
99--0.028--0.10--
1010.2990.3430.4030.260.2
1050.0110.0280.015----
13INU005 1060.0110.0560.0480.06--
1100.054--0.0410.10--
120----0.002----
1240.8240.7960.7880.561.0
1260.0500.0930.0320.08--
128----0.0110.02--
1300.018--0.0190.12--
1320.0430.0560.0580.04--
134------0.02--
14INU030 1440.2190.2500.2100.280.3
1460.0360.0280.032----
1480.0580.0830.0390.10--
1500.5250.5650.6540.420.5
1520.0830.0280.0110.080.2
1540.0790.0460.0540.10--
156------0.02--
15INU055 208------0.04--
2100.5250.5090.6080.460.3
2120.0290.0460.0280.06--
2140.1760.1300.1150.20--
2160.0650.0930.0710.02--
2180.2050.2220.1770.220.7
16LEI004 850.0760.1020.1100.18--
950.8270.8060.7450.700.8
970.0900.0930.1430.02--
101------0.02--
1050.004------0.2
1070.004--0.0020.08--
17REN105L03 2290.011--------
2310.0220.0830.0350.08--
2330.3270.2310.3070.08--
2350.1080.1300.1430.08--
2370.3060.2310.2680.24--
2410.2270.3240.2470.501.0
247------0.02--
18REN162C04 1980.004----0.14--
2000.0250.0460.0560.06--
2020.0970.1020.0910.120.3
2040.0500.0740.0580.040.3
2060.3170.2780.2340.060.3
2080.0360.0190.0500.06--
2100.4710.4810.5110.52--
19REN169D01 2020.2050.2040.1620.160.2
210--0.0190.0130.08--
2120.3490.3700.4180.180.2
2160.2590.2870.2790.420.7
2200.1870.1200.1280.16--
20REN169O18 158------0.04--
1600.043--0.0300.16--
1620.1730.2220.1360.40--
1640.6440.4810.6840.301.0
1660.0940.1850.102----
1680.0250.0280.0390.02--
1700.0220.0830.0090.06--
172------0.02--
21REN247M23 2680.2660.2410.2010.24--
2700.0220.0560.0370.22--
2720.3990.4260.4130.320.2
276--------0.3
2780.3130.2780.3480.220.5
22REN54P11 2220.0140.1020.0280.04--
2240.007--0.006----
2260.2050.2130.2320.060.2
230------0.02--
2320.2450.1760.1690.340.2
2340.0970.0370.0870.020.3
2360.2660.3430.3380.500.3
2380.1650.1300.1410.02--
23REN64E19 1390.1760.2410.1390.220.7
1430.014--0.009----
1450.4030.2690.4760.06--
1470.3710.4630.3140.56--
1490.0180.0190.0300.040.3
1510.007--0.0060.02--
1530.007--0.0260.08--
1550.0040.009--0.02--
24VGL0760 120.2520.4440.2710.140.3
130.0110.0370.011----
140.0400.0280.037----
20.20.0290.0190.0390.040.3
21.20.3240.2500.3270.36--
22.20.0320.0280.0740.14--
23.20.1510.0740.1300.24--
24.20.1580.1200.1100.080.3
25.20.004--0.002----
25VGL0910 15.1------0.02--
16.10.0650.0280.1210.16--
17.10.004--0.0040.08--
18.10.0140.0090.0370.04--
190.0830.0650.030----
19.10.2190.3060.2730.100.3
200.0250.056--0.06--
20.10.0110.0090.0220.160.2
210.004--------
21.10.1800.1300.2030.080.3
220.0320.0190.048----
22.10.3530.3610.2530.100.2
23----0.002----
23.10.007--0.0060.14--
24.10.0040.019--0.06--
26VGL1063 8----0.0020.02--
9------0.02--
11------0.02--
120.022--0.022----
130.022--0.0150.04--
140.2590.1480.2710.360.2
150.3960.4540.4070.320.8
160.1220.1670.1210.02--
170.0470.0650.0500.10--
180.0900.1390.0670.10--
190.0070.0090.002----
200.0360.0190.039----
21----0.004----
27VGL1165 15------0.02--
18------0.02--
190.4820.4540.5650.320.8
200.2480.1570.1100.14--
210.0790.0650.0560.060.2
220.004--------
250.0040.019--0.08--
260.0220.0740.0320.06--
27----0.004----
280.0860.0740.0840.26--
290.0720.1480.1300.04--
300.0040.0090.013----
31----0.004----
28VGL1828 140.1190.1110.082----
150.0430.0830.026--0.2
160.2050.2410.1600.280.3
170.1190.1480.1430.120.3
18--0.0090.0220.040.2
190.3710.2500.3610.26--
200.0470.0190.0190.02--
210.047--0.0500.14--
220.0500.1390.1340.10--
23----0.0020.04--
29VGL2009 100.011--0.0090.04--
110.2450.1940.2750.12--
120.029--0.0020.020.2
130.5720.6480.5190.440.3
140.0250.0460.0800.260.5
150.1080.1110.1000.12--
160.011--0.015----
30VGL2409 130.0140.0190.0040.020.2
140.007--0.0130.02--
150.007--0.0060.06--
160.0830.0830.1540.08--
170.2300.3150.2270.400.2
180.3130.2310.3380.380.7
190.011--0.0130.02--
200.3350.3520.2380.02--
21----0.006----
31VGL2918 120.0790.1760.1210.140.2
130.1220.0090.1490.140.2
140.0760.1110.0780.12--
150.022--0.0300.10--
160.0040.0090.015----
17.30.5360.4630.4650.280.7
18------0.02--
18.30.0860.1110.0320.14--
19.30.0760.1020.0950.06--
20.3--0.019------
22.3----0.013----
32VGL3008 130.0540.0560.028----
150.1330.0930.1580.140.3
160.0290.1020.0650.02--
170.1260.2040.0710.080.2
180.4240.2780.3480.320.2
190.0760.0280.1000.16--
200.1580.2410.2160.240.3
21----0.002----
22----0.0110.04--
33VGL3235 120.0040.0090.0130.04--
130.1220.0650.1060.04--
140.2300.2780.1950.380.2
150.1080.0930.0910.06--
160.0430.0370.0430.12--
170.2590.1390.2290.100.3
180.1580.1850.1860.160.5
190.0040.0090.0220.04--
200.0220.0090.0110.02--
210.0500.1760.1040.04--

Standard genetic assessment based on 33 autosomal STR loci

Biewer

  NNaNeHoHeF
Mean4528.3643.5220.6740.6850.017
 SE 0.4170.1840.0200.0190.005

Biewer Yorshire Terrier

  NNaNeHoHeF
Mean546.0003.3880.6900.672-0.029
 SE 0.2870.1730.0230.0210.014

Biewer Terrier

  NNaNeHoHeF
Mean2317.2733.4110.6590.6730.021
 SE 0.3670.1880.0210.0200.008

Yorkshire Terrier

  NNaNeHoHeF
Mean256.8483.8950.6950.7200.034
 SE 0.2910.2270.0180.0140.019

Biro Biewer

  NNaNeHoHeF
Mean32.6362.2110.5860.483-0.190
 SE 0.1470.1360.0570.0350.076

Standard genetic assessment based on 7 STRs in the DLA region

Biewer

  NNaNeHoHeF
Mean4526.7142.5700.5990.597-0.002
 SE 0.8010.1700.0300.0300.005

Biewer Yorshire Terrier

  NNaNeHoHeF
Mean546.1433.0840.6960.660-0.060
 SE 0.6520.2440.0260.0300.027

Biewer Terrier

  NNaNeHoHeF
Mean2315.2862.3680.5710.565-0.008
 SE 0.6910.1480.0340.0300.009

Yorkshire Terrier

  NNaNeHoHeF
Mean255.5712.8370.5200.6300.183
 SE 0.4450.2180.0540.0320.066

Biro Biewer

  NNaNeHoHeF
Mean32.4292.0960.7620.460-0.535
 SE 0.2750.2310.1460.0830.116

Standard genetic assessment for individual STR loci

Biewer

#LocusNNaNeHoHeF
1AHT121452124.9640.8140.799-0.020
2AHT137452104.2170.7880.763-0.032
3AHTH13045293.5690.7080.7200.016
4AHTh171-A45291.9880.4960.4970.003
5AHTh26045283.0990.6640.6770.020
6AHTk21145252.5130.5940.6020.013
7AHTk25345262.9490.6550.6610.009
8C22.27945283.8740.7350.7420.010
9FH200145282.4380.5730.5900.028
10FH205445264.1800.7630.761-0.003
11FH284845273.4360.7170.709-0.011
12INRA2145263.9170.7170.7450.037
13INU00545291.5850.3520.3690.047
14INU03045272.4610.6020.594-0.014
15INU05545262.6420.6060.6220.025
16LEI00445261.6010.3740.3760.004
17REN105L0345274.0370.7540.752-0.003
18REN162C0445273.0820.6460.6760.044
19REN169D0145253.6190.7210.7240.003
20REN169O1845282.2840.5460.5620.028
21REN247M2345253.1180.6920.679-0.019
22REN54P1145284.5740.7150.7810.085
23REN64E1945283.1010.6130.6780.095
24VGL076045294.5880.7280.7820.069
25VGL0910452155.1770.8190.807-0.015
26VGL1063452133.8800.7170.7420.034
27VGL1165452133.1640.6640.6840.030
28VGL1828452105.0730.8010.8030.002
29VGL200945272.6730.6080.6260.028
30VGL240945293.9820.7520.749-0.004
31VGL2918452113.6130.6920.7230.043
32VGL300845294.6540.7880.785-0.003
33VGL3235452106.1750.8160.8380.026

Biewer Yorshire Terrier

#LocusNNaNeHoHeF
1AHT1215473.8800.7960.742-0.073
2AHT1375464.1130.7590.757-0.003
3AHTH1305463.7870.7590.736-0.032
4AHTh171-A5441.5970.3700.3740.009
5AHTh2605472.7890.6110.6410.047
6AHTk2115442.0490.5000.5120.023
7AHTk2535453.1830.6480.6860.055
8C22.2795473.6400.7040.7250.030
9FH20015462.5600.5740.6090.058
10FH20545463.4210.6670.7080.058
11FH28485453.3730.7780.704-0.106
12INRA215464.0670.7040.7540.067
13INU0055441.5410.3890.351-0.107
14INU0305462.5500.5930.6080.025
15INU0555452.9740.6480.6640.024
16LEI0045431.4970.3520.332-0.059
17REN105L035454.2380.6670.7640.127
18REN162C045463.0550.7410.673-0.101
19REN169D015453.6250.8520.724-0.176
20REN169O185453.0940.7040.677-0.040
21REN247M235443.1290.7960.680-0.170
22REN54P115464.5000.7410.7780.048
23REN64E195452.9000.7590.655-0.159
24VGL07605483.5300.7960.717-0.111
25VGL091054104.0080.7780.751-0.036
26VGL10635473.5780.7590.721-0.054
27VGL11655483.7290.7220.7320.013
28VGL18285485.5120.8150.8190.005
29VGL20095442.1170.5930.528-0.123
30VGL24095453.5240.6850.7160.043
31VGL29185483.5600.8330.719-0.159
32VGL30085475.0150.8330.801-0.041
33VGL323554105.6790.8520.824-0.034

Biewer Terrier

#LocusNNaNeHoHeF
1AHT121231124.6910.7920.787-0.007
2AHT13723174.1900.8050.761-0.058
3AHTH13023183.2250.6670.6900.034
4AHTh171-A23181.8660.4760.464-0.026
5AHTh26023172.9790.6540.6640.016
6AHTk21123152.6180.6150.6180.005
7AHTk25323152.7660.6410.639-0.003
8C22.27923183.5810.7060.7210.021
9FH200123152.2930.5190.5640.079
10FH205423164.2390.7710.764-0.008
11FH284823173.3970.6930.7060.018
12INRA2123153.6190.7140.7240.013
13INU00523181.5880.3460.3700.065
14INU03023162.0960.5240.523-0.002
15INU05523152.3780.5840.580-0.008
16LEI00423141.7040.4160.413-0.006
17REN105L0323154.0160.7580.751-0.009
18REN162C0423163.0040.6100.6670.085
19REN169D0123153.3860.6710.7050.048
20REN169O1823162.0030.4940.5010.014
21REN247M2323142.9920.6320.6660.051
22REN54P1123174.4590.7230.7760.068
23REN64E1923172.8890.5580.6540.146
24VGL076023194.5980.6970.7830.109
25VGL0910231115.0080.8140.800-0.017
26VGL1063231113.8060.7320.7370.008
27VGL116523192.7800.6450.640-0.007
28VGL1828231104.8660.8050.794-0.013
29VGL200923172.7620.6320.6380.009
30VGL240923194.0580.8010.754-0.063
31VGL291823193.6890.6670.7290.085
32VGL300823194.6860.7530.7870.042
33VGL3235231106.3220.8400.8420.002

Yorkshire Terrier

#LocusNNaNeHoHeF
1AHT1212596.5450.8800.847-0.039
2AHT1372573.9680.8000.748-0.070
3AHTH1302583.9810.6800.7490.092
4AHTh171-A2573.2980.8000.697-0.148
5AHTh2602563.8340.6800.7390.080
6AHTk2112552.1370.5420.532-0.018
7AHTk2532563.2890.8000.696-0.149
8C22.2792583.6550.6800.7260.064
9FH20012573.9430.6400.7460.143
10FH20542553.4530.7600.710-0.070
11FH28482543.5010.6000.7140.160
12INRA212543.3070.7200.698-0.032
13INU0052582.8540.6400.6500.015
14INU0302563.5510.6400.7180.109
15INU0552563.2720.7600.694-0.094
16LEI0042551.8880.3600.4700.235
17REN105L032563.0560.7200.673-0.070
18REN162C042573.1570.7200.683-0.054
19REN169D012553.7540.6800.7340.073
20REN169O182573.5510.7600.718-0.058
21REN247M232543.8940.7200.7430.031
22REN54P112572.6880.4800.6280.236
23REN64E192572.6710.7200.626-0.151
24VGL07602564.2660.6400.7660.164
25VGL091025118.8650.7600.8870.143
26VGL10632593.9180.7600.745-0.020
27VGL11652594.8640.6800.7940.144
28VGL18282585.1650.8000.8060.008
29VGL20092563.4250.6000.7080.153
30VGL24092583.1650.6000.6840.123
31VGL29182586.0390.7200.8340.137
32VGL30082574.6820.8400.786-0.068
33VGL323525104.8830.7600.7950.044

Biro Biewer

#LocusNNaNeHoHeF
1AHT121332.0000.6670.500-0.333
2AHT137321.8000.6670.444-0.500
3AHTH130332.0000.3330.5000.333
4AHTh171-A332.5710.3330.6110.455
5AHTh260332.5711.0000.611-0.636
6AHTk211321.8000.6670.444-0.500
7AHTk253321.8000.6670.444-0.500
8C22.279321.3850.3330.278-0.200
9FH2001332.0000.6670.500-0.333
10FH2054322.0001.0000.500-1.000
11FH2848322.0001.0000.500-1.000
12INRA21343.6001.0000.722-0.385
13INU005311.0000.0000.0000.000
14INU030332.5710.6670.611-0.091
15INU055321.8000.0000.4441.000
16LEI004321.3850.3330.278-0.200
17REN105L03311.0000.0000.0000.000
18REN162C04333.0000.6670.6670.000
19REN169D01332.0000.6670.500-0.333
20REN169O18311.0000.0000.0000.000
21REN247M23332.5711.0000.611-0.636
22REN54P11343.6000.6670.7220.077
23REN64E19321.8000.0000.4441.000
24VGL0760333.0000.6670.6670.000
25VGL0910343.6001.0000.722-0.385
26VGL1063321.3850.3330.278-0.200
27VGL1165321.3850.3330.278-0.200
28VGL1828343.6000.6670.7220.077
29VGL2009332.5710.6670.611-0.091
30VGL2409332.0000.6670.500-0.333
31VGL2918332.0000.6670.500-0.333
32VGL3008343.6001.0000.722-0.385
33VGL3235332.5711.0000.611-0.636

Standard genetic assessment for 7 STRs in the DLA region

Biewer

#LocusNNaNeHoHeF
1DLA I-3CCA45282.2240.5440.5500.011
2DLA I-4ACA45292.3990.5950.583-0.020
3DLA I-4BCT45263.2480.6950.692-0.004
4DLA1131452102.9240.6530.6580.008
55ACA45242.8070.6570.644-0.021
65ACT45252.6130.6080.6170.014
75BCA45251.7750.4380.437-0.003

Biewer Yorshire Terrier

#LocusNNaNeHoHeF
1DLA I-3CCA5482.0390.5930.510-0.163
2DLA I-4ACA5473.0610.7410.673-0.100
3DLA I-4BCT5453.4370.6670.7090.060
4DLA11315494.1100.7410.7570.021
55ACA5443.5070.7960.715-0.114
65ACT5453.0360.7220.671-0.077
75BCA5452.3960.6110.583-0.049

Biewer Terrier

#LocusNNaNeHoHeF
1DLA I-3CCA23182.1670.5240.5390.027
2DLA I-4ACA23152.1730.5540.540-0.026
3DLA I-4BCT23153.0010.6930.667-0.039
4DLA113123182.6050.6280.616-0.019
55ACA23142.5260.6060.604-0.003
65ACT23142.4460.6060.591-0.025
75BCA23131.6560.3850.3960.027

Yorkshire Terrier

#LocusNNaNeHoHeF
1DLA I-3CCA2563.4720.6000.7120.157
2DLA I-4ACA2582.7900.6000.6420.065
3DLA I-4BCT2553.3970.6000.7060.150
4DLA11312563.4060.6400.7060.094
55ACA2542.5880.6000.6140.022
65ACT2552.3630.2400.5770.584
75BCA2551.8410.3600.4570.212

Biro Biewer

#LocusNNaNeHoHeF
1DLA I-3CCA332.5711.0000.611-0.636
2DLA I-4ACA332.5711.0000.611-0.636
3DLA I-4BCT332.5711.0000.611-0.636
4DLA1131321.3850.3330.278-0.200
55ACA332.5711.0000.611-0.636
65ACT322.0001.0000.500-1.000
75BCA311.0000.0000.0000.000