Statistics and Breed-wide Allele Frequency - Biewer

Updated Jul 15, 2026

DLA Class I Haplotype Frequencies

DLA1 #STR typesBiewer (n=139)Biewer Yorshire Terrier (n=54)Biewer Terrier (n=233)Yorkshire Terrier (n=16)Biro Biewer (n=3)
1005389 371 277 1810.0250.0090.013----
1007380 372 281 1820.0040.009--0.03--
1008386 373 289 1820.007----0.06--
1012388 369 289 1880.2230.1570.2320.280.2
1014375 373 287 1780.0290.0090.0580.09--
1016382 371 277 1780.0220.0190.0110.03--
1019380 373 287 1850.0720.0830.0600.09--
1030380 373 293 1780.4570.3520.4940.310.5
1035386 373 277 184--0.009------
1040380 371 277 1860.0940.2040.077----
1043393 381 277 1830.0070.0090.011----
1046376 379 291 1800.029--0.021----
1068380 373 287 181--0.009------
1087380 371 277 1780.004--------
1092376 379 277 181--0.037--0.06--
1104386 373 289 1860.0110.0190.013----
1105382 379 277 1780.004------0.3
1115386 371 277 182----0.002----
1117376 373 277 180----0.002----
1118376 377 277 1800.0040.019------
1130380 373 287 1780.0070.009------
1131388 379 277 1810.0040.0460.0060.03--

DLA Class II Haplotype Frequencies

DLA2 #STR typesBiewer (n=139)Biewer Yorshire Terrier (n=54)Biewer Terrier (n=233)Yorkshire Terrier (n=16)Biro Biewer (n=3)
2003343 324 2820.2590.2130.2510.380.2
2005339 322 2800.0400.0090.021----
2013345 327 2840.0470.1390.064----
2016339 323 2840.0790.0930.0710.09--
2021339 324 2680.004--------
2023341 323 2820.4570.3520.4960.310.5
2037341 327 2800.0290.0090.0580.09--
2040345 327 2800.0610.0830.0240.03--
2044343 324 268--0.037--0.06--
2053343 324 280--0.009------
2074341 324 2840.0040.0190.002----
2080339 325 2760.0040.009--0.03--
2081343 322 2820.0110.0190.013----
2082339 325 268--0.009------
2083339 324 2820.004------0.3
2084339 323 2680.004--------

Allele Frequencies

#Locus NameAlleleBiewer (n=142)Biewer Yorshire Terrier (n=54)Biewer Terrier (n=233)Yorkshire Terrier (n=25)Biro Biewer (n=3)
1AHT121 92----0.0150.040.2
940.004--0.0110.06--
980.1060.1480.1050.20--
1000.2060.0650.1140.20--
1020.3010.3060.3540.120.7
1040.2130.3610.2210.08--
1060.032--0.0130.08--
1080.1060.0830.1240.20--
1100.0070.0280.0190.02--
1120.0250.0090.019----
122----0.002--0.2
124----0.002----
2AHT137 1310.2360.2780.2400.200.3
133------0.08--
1370.3270.2780.3130.40--
1390.1270.1200.109----
143--0.009------
147----0.0110.04--
1490.004--0.004----
1510.2430.2690.2550.20--
1530.0630.0460.0670.060.7
155------0.02--
3AHTH130 1170.011--0.011----
1190.3660.3890.3350.140.7
1210.1650.1570.0920.080.2
1230.0210.0740.0170.08--
125----0.0020.02--
1270.3380.2690.4210.240.2
1290.0920.1020.1050.40--
1310.0070.0090.0170.02--
133------0.02--
4AHTh171-A 217----0.002----
2190.6480.7780.7100.500.3
2250.1970.0930.1330.120.5
227----0.0020.04--
2290.004--0.0020.04--
2310.1090.1110.1160.120.2
2330.0280.0190.0280.14--
2350.011--0.006----
2370.004----0.04--
5AHTh260 2380.0460.0740.0210.22--
240------0.02--
2420.0420.0190.0260.12--
2440.0850.0370.0710.020.2
2460.2570.2410.2680.360.3
2480.0740.0650.114----
2500.4960.5370.4960.260.5
254--0.0280.004----
6AHTk211 870.4720.5460.4890.210.7
890.018--0.0210.04--
910.4120.4350.3690.650.3
950.0210.0090.0580.06--
970.0770.0090.0620.04--
7AHTk253 2840.0490.0090.0560.020.3
2860.0460.0280.0580.02--
2880.1940.3430.2170.42--
2900.1480.2690.1290.24--
2920.5530.3520.5410.260.7
2940.011----0.04--
8C22.279 1160.1130.1110.0900.020.2
1180.0740.0370.0360.12--
1200.0040.0090.0090.04--
1220.0110.0090.0090.04--
1240.1830.1940.1950.04--
1260.3350.4170.4310.460.8
1280.2640.2220.2190.18--
1300.018--0.0110.10--
9FH2001 124------0.12--
1320.4790.4720.5240.400.2
136--0.009--0.02--
140----0.0090.12--
1440.0850.0930.0520.080.2
1480.4230.3980.3990.240.7
1520.0140.0090.0170.02--
156--0.019------
10FH2054 1480.2220.2590.2080.34--
1520.1270.0560.1570.260.5
1560.3490.4440.3540.320.5
1600.2040.1390.195----
1640.0600.0460.0620.02--
1680.0390.0560.0240.06--
11FH2848 2320.011--0.002----
2340.0920.0560.077----
2360.3200.3330.3410.24--
2380.1730.1300.1570.220.5
2400.3870.3980.3820.400.5
242----0.002----
2440.0180.0830.0390.14--
12INRA21 910.1900.1940.210--0.2
950.2010.1480.2000.420.3
970.2890.2590.1720.220.3
99--0.028--0.10--
1010.3100.3430.4030.260.2
1050.0110.0280.015----
13INU005 1060.0110.0560.0470.06--
1100.063--0.0410.10--
120----0.002----
1240.8170.7960.7900.561.0
1260.0490.0930.0320.08--
128----0.0110.02--
1300.018--0.0190.12--
1320.0420.0560.0580.04--
134------0.02--
14INU030 1440.2180.2500.2080.280.3
1460.0350.0280.034----
1480.0560.0830.0390.10--
1500.5320.5650.6550.420.5
1520.0810.0280.0110.080.2
1540.0770.0460.0540.10--
156------0.02--
15INU055 208------0.04--
2100.5250.5090.6120.460.3
2120.0280.0460.0280.06--
2140.1800.1300.1140.20--
2160.0670.0930.0710.02--
2180.2010.2220.1760.220.7
16LEI004 850.0810.1020.1120.18--
950.8240.8060.7450.700.8
970.0880.0930.1420.02--
101------0.02--
1050.004------0.2
1070.004--0.0020.08--
17REN105L03 2290.011--------
2310.0210.0830.0340.08--
2330.3240.2310.3070.08--
2350.1230.1300.1420.08--
2370.2990.2310.2700.24--
2410.2220.3240.2470.501.0
247------0.02--
18REN162C04 1980.004----0.14--
2000.0250.0460.0560.06--
2020.0950.1020.0900.120.3
2040.0490.0740.0580.040.3
2060.3200.2780.2360.060.3
2080.0350.0190.0520.06--
2100.4720.4810.5090.52--
19REN169D01 2020.2080.2040.1610.160.2
210--0.0190.0130.08--
2120.3420.3700.4210.180.2
2160.2680.2870.2790.420.7
2200.1830.1200.1270.16--
20REN169O18 158------0.04--
1600.042--0.0300.16--
1620.1690.2220.1370.40--
1640.6440.4810.6820.301.0
1660.0920.1850.103----
1680.0320.0280.0390.02--
1700.0210.0830.0090.06--
172------0.02--
21REN247M23 2680.2750.2410.2000.24--
2700.0210.0560.0360.22--
2720.3910.4260.4140.320.2
276--------0.3
2780.3130.2780.3500.220.5
22REN54P11 2220.0140.1020.0300.04--
2240.007--0.006----
2260.2180.2130.2320.060.2
230------0.02--
2320.2430.1760.1700.340.2
2340.0950.0370.0860.020.3
2360.2610.3430.3350.500.3
2380.1620.1300.1420.02--
23REN64E19 1390.1730.2410.1390.220.7
1430.014--0.009----
1450.4080.2690.4760.06--
1470.3700.4630.3130.56--
1490.0180.0190.0300.040.3
1510.007--0.0060.02--
1530.007--0.0260.08--
1550.0040.009--0.02--
24VGL0760 120.2460.4440.2730.140.3
130.0110.0370.011----
140.0390.0280.039----
20.20.0280.0190.0390.040.3
21.20.3350.2500.3240.36--
22.20.0320.0280.0730.14--
23.20.1510.0740.1290.24--
24.20.1550.1200.1120.080.3
25.20.004--0.002----
25VGL0910 15.1------0.02--
16.10.0630.0280.1200.16--
17.10.004--0.0040.08--
18.10.0210.0090.0360.04--
190.0810.0650.030----
19.10.2150.3060.2730.100.3
200.0250.056--0.06--
20.10.0110.0090.0210.160.2
210.004--------
21.10.1870.1300.2020.080.3
220.0320.0190.049----
22.10.3490.3610.2550.100.2
23----0.002----
23.10.007--0.0060.14--
24.10.0040.019--0.06--
26VGL1063 8----0.0020.02--
9------0.02--
11------0.02--
120.021--0.021----
130.021--0.0150.04--
140.2540.1480.2730.360.2
150.3980.4540.4060.320.8
160.1270.1670.1220.02--
170.0460.0650.0490.10--
180.0920.1390.0670.10--
190.0070.0090.002----
200.0350.0190.039----
21----0.004----
27VGL1165 15------0.02--
18------0.02--
190.4820.4540.5640.320.8
200.2430.1570.1120.14--
210.0770.0650.0560.060.2
220.004--------
250.0040.019--0.08--
260.0250.0740.0320.06--
27----0.004----
280.0850.0740.0840.26--
290.0770.1480.1290.04--
300.0040.0090.015----
31----0.004----
28VGL1828 140.1270.1110.084----
150.0420.0830.026--0.2
160.2010.2410.1590.280.3
170.1160.1480.1440.120.3
18--0.0090.0210.040.2
190.3660.2500.3610.26--
200.0460.0190.0190.02--
210.046--0.0520.14--
220.0560.1390.1330.10--
23----0.0020.04--
29VGL2009 100.011--0.0090.04--
110.2430.1940.2770.12--
120.028--0.0020.020.2
130.5700.6480.5170.440.3
140.0250.0460.0790.260.5
150.1130.1110.1010.12--
160.011--0.015----
30VGL2409 130.0140.0190.0040.020.2
140.007--0.0130.02--
150.007--0.0060.06--
160.0920.0830.1520.08--
170.2320.3150.2250.400.2
180.3100.2310.3410.380.7
190.011--0.0130.02--
200.3270.3520.2380.02--
21----0.006----
31VGL2918 120.0770.1760.1200.140.2
130.1200.0090.1480.140.2
140.0740.1110.0770.12--
150.025--0.0300.10--
160.0040.0090.015----
17.30.5250.4630.4680.280.7
18------0.02--
18.30.0850.1110.0320.14--
19.30.0920.1020.0970.06--
20.3--0.019------
22.3----0.013----
32VGL3008 130.0530.0560.028----
150.1300.0930.1570.140.3
160.0350.1020.0640.02--
170.1230.2040.0730.080.2
180.4190.2780.3480.320.2
190.0740.0280.1030.16--
200.1650.2410.2150.240.3
21----0.002----
22----0.0110.04--
33VGL3235 120.0040.0090.0130.04--
130.1200.0650.1050.04--
140.2360.2780.1950.380.2
150.1060.0930.0900.06--
160.0420.0370.0430.12--
170.2570.1390.2300.100.3
180.1550.1850.1870.160.5
190.0040.0090.0210.04--
200.0280.0090.0110.02--
210.0490.1760.1050.04--

Standard genetic assessment based on 33 autosomal STR loci

Biewer

  NNaNeHoHeF
Mean4578.3643.5220.6740.6850.017
 SE 0.4170.1850.0200.0190.005

Biewer Yorshire Terrier

  NNaNeHoHeF
Mean546.0003.3880.6900.672-0.029
 SE 0.2870.1730.0230.0210.014

Biewer Terrier

  NNaNeHoHeF
Mean2337.2733.4080.6590.6730.021
 SE 0.3670.1890.0210.0200.008

Yorkshire Terrier

  NNaNeHoHeF
Mean256.8483.8950.6950.7200.034
 SE 0.2910.2270.0180.0140.019

Biro Biewer

  NNaNeHoHeF
Mean32.6362.2110.5860.483-0.190
 SE 0.1470.1360.0570.0350.076

Standard genetic assessment based on 7 STRs in the DLA region

Biewer

  NNaNeHoHeF
Mean4576.7142.5630.5960.5960.001
 SE 0.8010.1700.0300.0300.005

Biewer Yorshire Terrier

  NNaNeHoHeF
Mean546.1433.0840.6960.660-0.060
 SE 0.6520.2440.0260.0300.027

Biewer Terrier

  NNaNeHoHeF
Mean2335.2862.3740.5700.566-0.005
 SE 0.6910.1490.0340.0310.009

Yorkshire Terrier

  NNaNeHoHeF
Mean255.5712.8370.5200.6300.183
 SE 0.4450.2180.0540.0320.066

Biro Biewer

  NNaNeHoHeF
Mean32.4292.0960.7620.460-0.535
 SE 0.2750.2310.1460.0830.116

Standard genetic assessment for individual STR loci

Biewer

#LocusNNaNeHoHeF
1AHT121457124.9450.8140.798-0.020
2AHT137457104.2260.7900.763-0.035
3AHTH13045793.5670.7090.7200.015
4AHTh171-A45791.9780.4920.4940.004
5AHTh26045783.0740.6590.6750.024
6AHTk21145752.5210.5960.6030.011
7AHTk25345762.9270.6500.6580.013
8C22.27945783.8280.7260.7390.017
9FH200145782.4440.5780.5910.022
10FH205445764.1910.7640.761-0.003
11FH284845773.4410.7180.709-0.012
12INRA2145763.8970.7110.7430.043
13INU00545791.5890.3540.3710.043
14INU03045772.4490.6000.592-0.013
15INU05545762.6350.6040.6200.027
16LEI00445761.6040.3760.3770.001
17REN105L0345774.0530.7530.7530.001
18REN162C0445773.0810.6500.6750.038
19REN169D0145753.6150.7200.7230.005
20REN169O1845782.2870.5490.5630.024
21REN247M2345753.1220.6940.680-0.021
22REN54P1145784.5890.7130.7820.088
23REN64E1945783.0910.6130.6770.094
24VGL076045794.5760.7260.7810.070
25VGL0910457155.1870.8210.807-0.017
26VGL1063457133.8800.7180.7420.033
27VGL1165457133.1680.6650.6840.028
28VGL1828457105.1020.8030.8040.001
29VGL200945772.6810.6130.6270.023
30VGL240945793.9860.7530.749-0.005
31VGL2918457113.6350.6890.7250.049
32VGL300845794.6740.7880.786-0.002
33VGL3235457106.1720.8180.8380.023

Biewer Yorshire Terrier

#LocusNNaNeHoHeF
1AHT1215473.8800.7960.742-0.073
2AHT1375464.1130.7590.757-0.003
3AHTH1305463.7870.7590.736-0.032
4AHTh171-A5441.5970.3700.3740.009
5AHTh2605472.7890.6110.6410.047
6AHTk2115442.0490.5000.5120.023
7AHTk2535453.1830.6480.6860.055
8C22.2795473.6400.7040.7250.030
9FH20015462.5600.5740.6090.058
10FH20545463.4210.6670.7080.058
11FH28485453.3730.7780.704-0.106
12INRA215464.0670.7040.7540.067
13INU0055441.5410.3890.351-0.107
14INU0305462.5500.5930.6080.025
15INU0555452.9740.6480.6640.024
16LEI0045431.4970.3520.332-0.059
17REN105L035454.2380.6670.7640.127
18REN162C045463.0550.7410.673-0.101
19REN169D015453.6250.8520.724-0.176
20REN169O185453.0940.7040.677-0.040
21REN247M235443.1290.7960.680-0.170
22REN54P115464.5000.7410.7780.048
23REN64E195452.9000.7590.655-0.159
24VGL07605483.5300.7960.717-0.111
25VGL091054104.0080.7780.751-0.036
26VGL10635473.5780.7590.721-0.054
27VGL11655483.7290.7220.7320.013
28VGL18285485.5120.8150.8190.005
29VGL20095442.1170.5930.528-0.123
30VGL24095453.5240.6850.7160.043
31VGL29185483.5600.8330.719-0.159
32VGL30085475.0150.8330.801-0.041
33VGL323554105.6790.8520.824-0.034

Biewer Terrier

#LocusNNaNeHoHeF
1AHT121233124.6520.7900.785-0.006
2AHT13723374.2070.8070.762-0.058
3AHTH13023383.2340.6650.6910.037
4AHTh171-A23381.8640.4760.464-0.028
5AHTh26023372.9690.6520.6630.016
6AHTk21123352.6090.6140.6170.005
7AHTk25323352.7590.6390.638-0.003
8C22.27923383.5490.7000.7180.026
9FH200123352.2910.5240.5640.071
10FH205423364.2410.7730.764-0.011
11FH284823373.3980.6910.7060.021
12INRA2123353.6160.7080.7230.021
13INU00523381.5810.3430.3680.066
14INU03023362.0950.5240.523-0.002
15INU05523352.3600.5790.576-0.005
16LEI00423341.7040.4160.413-0.008
17REN105L0323354.0090.7600.751-0.012
18REN162C0423363.0150.6140.6680.082
19REN169D0123353.3690.6700.7030.048
20REN169O1823362.0100.4980.5020.009
21REN247M2323342.9850.6350.6650.045
22REN54P1123374.4870.7250.7770.067
23REN64E1923372.8880.5580.6540.147
24VGL076023394.6140.7000.7830.107
25VGL0910233115.0050.8150.800-0.019
26VGL1063233113.8050.7300.7370.010
27VGL116523392.7860.6440.641-0.004
28VGL1828233104.8850.8070.795-0.015
29VGL200923372.7710.6350.6390.006
30VGL240923394.0390.7980.752-0.061
31VGL291823393.6650.6650.7270.085
32VGL300823394.7080.7510.7880.046
33VGL3235233106.3080.8410.8410.000

Yorkshire Terrier

#LocusNNaNeHoHeF
1AHT1212596.5450.8800.847-0.039
2AHT1372573.9680.8000.748-0.070
3AHTH1302583.9810.6800.7490.092
4AHTh171-A2573.2980.8000.697-0.148
5AHTh2602563.8340.6800.7390.080
6AHTk2112552.1370.5420.532-0.018
7AHTk2532563.2890.8000.696-0.149
8C22.2792583.6550.6800.7260.064
9FH20012573.9430.6400.7460.143
10FH20542553.4530.7600.710-0.070
11FH28482543.5010.6000.7140.160
12INRA212543.3070.7200.698-0.032
13INU0052582.8540.6400.6500.015
14INU0302563.5510.6400.7180.109
15INU0552563.2720.7600.694-0.094
16LEI0042551.8880.3600.4700.235
17REN105L032563.0560.7200.673-0.070
18REN162C042573.1570.7200.683-0.054
19REN169D012553.7540.6800.7340.073
20REN169O182573.5510.7600.718-0.058
21REN247M232543.8940.7200.7430.031
22REN54P112572.6880.4800.6280.236
23REN64E192572.6710.7200.626-0.151
24VGL07602564.2660.6400.7660.164
25VGL091025118.8650.7600.8870.143
26VGL10632593.9180.7600.745-0.020
27VGL11652594.8640.6800.7940.144
28VGL18282585.1650.8000.8060.008
29VGL20092563.4250.6000.7080.153
30VGL24092583.1650.6000.6840.123
31VGL29182586.0390.7200.8340.137
32VGL30082574.6820.8400.786-0.068
33VGL323525104.8830.7600.7950.044

Biro Biewer

#LocusNNaNeHoHeF
1AHT121332.0000.6670.500-0.333
2AHT137321.8000.6670.444-0.500
3AHTH130332.0000.3330.5000.333
4AHTh171-A332.5710.3330.6110.455
5AHTh260332.5711.0000.611-0.636
6AHTk211321.8000.6670.444-0.500
7AHTk253321.8000.6670.444-0.500
8C22.279321.3850.3330.278-0.200
9FH2001332.0000.6670.500-0.333
10FH2054322.0001.0000.500-1.000
11FH2848322.0001.0000.500-1.000
12INRA21343.6001.0000.722-0.385
13INU005311.0000.0000.0000.000
14INU030332.5710.6670.611-0.091
15INU055321.8000.0000.4441.000
16LEI004321.3850.3330.278-0.200
17REN105L03311.0000.0000.0000.000
18REN162C04333.0000.6670.6670.000
19REN169D01332.0000.6670.500-0.333
20REN169O18311.0000.0000.0000.000
21REN247M23332.5711.0000.611-0.636
22REN54P11343.6000.6670.7220.077
23REN64E19321.8000.0000.4441.000
24VGL0760333.0000.6670.6670.000
25VGL0910343.6001.0000.722-0.385
26VGL1063321.3850.3330.278-0.200
27VGL1165321.3850.3330.278-0.200
28VGL1828343.6000.6670.7220.077
29VGL2009332.5710.6670.611-0.091
30VGL2409332.0000.6670.500-0.333
31VGL2918332.0000.6670.500-0.333
32VGL3008343.6001.0000.722-0.385
33VGL3235332.5711.0000.611-0.636

Standard genetic assessment for 7 STRs in the DLA region

Biewer

#LocusNNaNeHoHeF
1DLA I-3CCA45782.2180.5400.5490.016
2DLA I-4ACA45792.4050.5950.584-0.018
3DLA I-4BCT45763.2360.6910.691-0.001
4DLA1131457102.9150.6500.6570.011
55ACA45742.7990.6540.643-0.018
65ACT45752.6020.6040.6160.019
75BCA45751.7660.4340.434-0.001

Biewer Yorshire Terrier

#LocusNNaNeHoHeF
1DLA I-3CCA5482.0390.5930.510-0.163
2DLA I-4ACA5473.0610.7410.673-0.100
3DLA I-4BCT5453.4370.6670.7090.060
4DLA11315494.1100.7410.7570.021
55ACA5443.5070.7960.715-0.114
65ACT5453.0360.7220.671-0.077
75BCA5452.3960.6110.583-0.049

Biewer Terrier

#LocusNNaNeHoHeF
1DLA I-3CCA23382.1770.5240.5410.032
2DLA I-4ACA23352.1840.5540.542-0.021
3DLA I-4BCT23353.0050.6910.667-0.036
4DLA113123382.6140.6270.617-0.015
55ACA23342.5330.6050.6050.000
65ACT23342.4540.6050.592-0.021
75BCA23331.6490.3820.3930.029

Yorkshire Terrier

#LocusNNaNeHoHeF
1DLA I-3CCA2563.4720.6000.7120.157
2DLA I-4ACA2582.7900.6000.6420.065
3DLA I-4BCT2553.3970.6000.7060.150
4DLA11312563.4060.6400.7060.094
55ACA2542.5880.6000.6140.022
65ACT2552.3630.2400.5770.584
75BCA2551.8410.3600.4570.212

Biro Biewer

#LocusNNaNeHoHeF
1DLA I-3CCA332.5711.0000.611-0.636
2DLA I-4ACA332.5711.0000.611-0.636
3DLA I-4BCT332.5711.0000.611-0.636
4DLA1131321.3850.3330.278-0.200
55ACA332.5711.0000.611-0.636
65ACT322.0001.0000.500-1.000
75BCA311.0000.0000.0000.000