Updated Aug 27, 2026
DLA Class I Haplotype Frequencies
| DLA1 # | STR types | Havanese (n=1175) |
|---|---|---|
| 1003 | 387 375 277 186 | 0.0387 |
| 1006 | 387 375 293 180 | 0.0549 |
| 1009 | 382 377 277 184 | 0.0004 |
| 1011 | 376 365 281 180 | 0.0009 |
| 1012 | 388 369 289 188 | 0.0111 |
| 1014 | 375 373 287 178 | 0.0183 |
| 1016 | 382 371 277 178 | 0.1923 |
| 1017 | 386 373 289 178 | 0.0009 |
| 1018 | 375 373 287 186 | 0.0068 |
| 1019 | 380 373 287 185 | 0.0009 |
| 1029 | 380 365 281 182 | 0.0357 |
| 1030 | 380 373 293 178 | 0.0026 |
| 1033 | 382 379 277 181 | 0.0013 |
| 1035 | 386 373 277 184 | 0.0038 |
| 1040 | 380 371 277 186 | 0.0157 |
| 1052 | 380 372 289 184 | 0.0089 |
| 1054 | 382 379 277 184 | 0.1230 |
| 1062 | 382 371 277 183 | 0.0004 |
| 1068 | 380 373 287 181 | 0.0128 |
| 1071 | 380 373 277 178 | 0.0004 |
| 1084 | 376 373 277 184 | 0.0013 |
| 1087 | 380 371 277 178 | 0.0034 |
| 1092 | 376 379 277 181 | 0.0962 |
| 1093 | 386 379 277 180 | 0.0136 |
| 1105 | 382 379 277 178 | 0.0009 |
| 1111 | 387 378 287 182 | 0.0026 |
| 1114 | 380 373 287 183 | 0.0170 |
| 1115 | 386 371 277 182 | 0.0923 |
| 1116 | 380 365 289 186 | 0.1081 |
| 1117 | 376 373 277 180 | 0.0477 |
| 1118 | 376 377 277 180 | 0.0009 |
| 1120 | 376 386 289 176 | 0.0009 |
| 1121 | 380 371 277 183 | 0.0047 |
| 1123 | 386 379 277 184 | 0.0013 |
| 1124 | 388 373 289 178 | 0.0055 |
| 1125 | 393 383 277 185 | 0.0217 |
| 1128 | 384 376 287 182 | 0.0009 |
| 1129 | 382 371 277 181 | 0.0009 |
| 1132 | 376 379 277 184 | 0.0013 |
| 1133 | 378 365 287 172 | 0.0170 |
| 1134 | 384 365 291 178 | 0.0009 |
| 1136 | 382 371 277 182 | 0.0004 |
| 1140 | 376 379 301 180 | 0.0038 |
| 1142 | 376 379 277 180 | 0.0047 |
| 1147 | 391 375 293 180 | 0.0026 |
| 1148 | 376 375 277 180 | 0.0021 |
| 1154 | 376 365 281 183 | 0.0098 |
| 1155 | 388 369 287 184 | 0.0004 |
| 1173 | 392 371 277 186 | 0.0038 |
| 1180 | 386 371 277 181 | 0.0004 |
| 1260 | 387 379 277 184 | 0.0004 |
| 1262 | 382 377 289 180 | 0.0009 |
| 1266 | 378 365 287 181 | 0.0009 |
| 1290 | 380 369 277 181 | 0.0009 |
| 1300 | 388 369 260 184 | 0.0004 |
DLA Class II Haplotype Frequencies
| DLA2 # | STR types | Havanese (n=1175) |
|---|---|---|
| 2001 | 343 324 284 | 0.0430 |
| 2003 | 343 324 282 | 0.2140 |
| 2005 | 339 322 280 | 0.0026 |
| 2006 | 339 325 280 | 0.0013 |
| 2007 | 351 327 280 | 0.0574 |
| 2008 | 339 327 276 | 0.0004 |
| 2011 | 345 322 284 | 0.0004 |
| 2012 | 345 322 280 | 0.0026 |
| 2014 | 339 322 284 | 0.0034 |
| 2016 | 339 323 284 | 0.0264 |
| 2017 | 343 322 280 | 0.0098 |
| 2018 | 339 324 284 | 0.0251 |
| 2021 | 339 324 268 | 0.0047 |
| 2022 | 339 327 282 | 0.1157 |
| 2023 | 341 323 282 | 0.0026 |
| 2024 | 343 323 280 | 0.0034 |
| 2028 | 345 327 288 | 0.0009 |
| 2032 | 339 323 280 | 0.0183 |
| 2033 | 339 323 282 | 0.0013 |
| 2035 | 341 323 280 | 0.0021 |
| 2037 | 341 327 280 | 0.0183 |
| 2040 | 345 327 280 | 0.0038 |
| 2053 | 343 324 280 | 0.0285 |
| 2066 | 339 324 280 | 0.1638 |
| 2068 | 339 327 284 | 0.0004 |
| 2070 | 347 324 282 | 0.1077 |
| 2071 | 339 322 286 | 0.0047 |
| 2072 | 339 325 282 | 0.0170 |
| 2073 | 339 327 286 | 0.0085 |
| 2074 | 341 324 284 | 0.0477 |
| 2075 | 341 327 282 | 0.0366 |
| 2076 | 345 322 282 | 0.0055 |
| 2077 | 347 325 286 | 0.0170 |
| 2079 | 343 323 278 | 0.0009 |
| 2082 | 339 325 268 | 0.0038 |
| 2087 | 347 324 280 | 0.0004 |
Allele Frequencies
| # | Locus Name | Allele | Havanese (n=1191) |
|---|---|---|---|
| 1 | AHT121 | 88 | 0.0004 |
| 92 | 0.1062 | ||
| 94 | 0.0764 | ||
| 96 | 0.0021 | ||
| 98 | 0.0097 | ||
| 100 | 0.0470 | ||
| 102 | 0.1058 | ||
| 104 | 0.5311 | ||
| 106 | 0.0348 | ||
| 108 | 0.0823 | ||
| 110 | 0.0013 | ||
| 112 | 0.0029 | ||
| 2 | AHT137 | 131 | 0.2334 |
| 133 | 0.0617 | ||
| 137 | 0.0819 | ||
| 139 | 0.0399 | ||
| 141 | 0.1788 | ||
| 143 | 0.1117 | ||
| 145 | 0.0080 | ||
| 147 | 0.2003 | ||
| 149 | 0.0642 | ||
| 151 | 0.0193 | ||
| 155 | 0.0008 | ||
| 3 | AHTH130 | 117 | 0.0361 |
| 119 | 0.2662 | ||
| 121 | 0.0495 | ||
| 123 | 0.0311 | ||
| 125 | 0.3669 | ||
| 127 | 0.1864 | ||
| 129 | 0.0407 | ||
| 131 | 0.0202 | ||
| 133 | 0.0029 | ||
| 4 | AHTh171-A | 217 | 0.0017 |
| 219 | 0.1608 | ||
| 221 | 0.0013 | ||
| 225 | 0.2552 | ||
| 227 | 0.0294 | ||
| 229 | 0.1864 | ||
| 231 | 0.2779 | ||
| 233 | 0.0206 | ||
| 235 | 0.0386 | ||
| 237 | 0.0281 | ||
| 5 | AHTh260 | 238 | 0.1243 |
| 242 | 0.0134 | ||
| 244 | 0.2343 | ||
| 246 | 0.2876 | ||
| 248 | 0.2040 | ||
| 250 | 0.1050 | ||
| 252 | 0.0025 | ||
| 254 | 0.0273 | ||
| 258 | 0.0017 | ||
| 6 | AHTk211 | 87 | 0.4740 |
| 89 | 0.2254 | ||
| 91 | 0.1465 | ||
| 95 | 0.1432 | ||
| 97 | 0.0109 | ||
| 7 | AHTk253 | 284 | 0.0118 |
| 286 | 0.0697 | ||
| 288 | 0.6432 | ||
| 290 | 0.0487 | ||
| 292 | 0.2267 | ||
| 8 | C22.279 | 116 | 0.1037 |
| 118 | 0.2128 | ||
| 120 | 0.0050 | ||
| 122 | 0.0168 | ||
| 124 | 0.2288 | ||
| 126 | 0.1327 | ||
| 128 | 0.0008 | ||
| 130 | 0.2993 | ||
| 9 | FH2001 | 124 | 0.0403 |
| 128 | 0.0088 | ||
| 132 | 0.4127 | ||
| 136 | 0.0113 | ||
| 140 | 0.0109 | ||
| 144 | 0.4530 | ||
| 148 | 0.0516 | ||
| 152 | 0.0097 | ||
| 158 | 0.0017 | ||
| 10 | FH2054 | 148 | 0.0160 |
| 152 | 0.0844 | ||
| 156 | 0.2510 | ||
| 160 | 0.1986 | ||
| 164 | 0.0882 | ||
| 168 | 0.1465 | ||
| 172 | 0.1100 | ||
| 176 | 0.1020 | ||
| 180 | 0.0029 | ||
| 184 | 0.0004 | ||
| 11 | FH2848 | 230 | 0.0139 |
| 232 | 0.3044 | ||
| 234 | 0.0487 | ||
| 236 | 0.0823 | ||
| 238 | 0.0466 | ||
| 240 | 0.2343 | ||
| 242 | 0.2603 | ||
| 244 | 0.0055 | ||
| 246 | 0.0042 | ||
| 12 | INRA21 | 91 | 0.1373 |
| 95 | 0.0995 | ||
| 97 | 0.3749 | ||
| 99 | 0.0504 | ||
| 101 | 0.1776 | ||
| 103 | 0.1495 | ||
| 105 | 0.0109 | ||
| 13 | INU005 | 106 | 0.0008 |
| 110 | 0.0306 | ||
| 122 | 0.0134 | ||
| 124 | 0.5882 | ||
| 126 | 0.2175 | ||
| 128 | 0.0088 | ||
| 132 | 0.0273 | ||
| 134 | 0.0038 | ||
| 138 | 0.1096 | ||
| 14 | INU030 | 144 | 0.6343 |
| 146 | 0.0386 | ||
| 148 | 0.0235 | ||
| 150 | 0.1276 | ||
| 152 | 0.1612 | ||
| 154 | 0.0134 | ||
| 156 | 0.0013 | ||
| 15 | INU055 | 210 | 0.3736 |
| 212 | 0.2061 | ||
| 214 | 0.1003 | ||
| 216 | 0.1297 | ||
| 218 | 0.1847 | ||
| 222 | 0.0055 | ||
| 16 | LEI004 | 83 | 0.0004 |
| 85 | 0.6419 | ||
| 93 | 0.0004 | ||
| 95 | 0.2254 | ||
| 97 | 0.0134 | ||
| 101 | 0.0017 | ||
| 105 | 0.0004 | ||
| 107 | 0.1092 | ||
| 109 | 0.0071 | ||
| 17 | REN105L03 | 227 | 0.0223 |
| 229 | 0.0181 | ||
| 231 | 0.2091 | ||
| 233 | 0.0924 | ||
| 235 | 0.0034 | ||
| 237 | 0.1906 | ||
| 239 | 0.0113 | ||
| 241 | 0.4471 | ||
| 245 | 0.0059 | ||
| 18 | REN162C04 | 200 | 0.0348 |
| 202 | 0.2200 | ||
| 204 | 0.0491 | ||
| 206 | 0.3967 | ||
| 208 | 0.1285 | ||
| 210 | 0.1709 | ||
| 19 | REN169D01 | 202 | 0.0684 |
| 204 | 0.0008 | ||
| 210 | 0.1683 | ||
| 212 | 0.0516 | ||
| 214 | 0.0269 | ||
| 216 | 0.4215 | ||
| 218 | 0.0701 | ||
| 220 | 0.1923 | ||
| 20 | REN169O18 | 160 | 0.0697 |
| 162 | 0.2045 | ||
| 164 | 0.0424 | ||
| 166 | 0.2322 | ||
| 168 | 0.1860 | ||
| 170 | 0.2347 | ||
| 172 | 0.0306 | ||
| 21 | REN247M23 | 266 | 0.0004 |
| 268 | 0.4110 | ||
| 270 | 0.1037 | ||
| 272 | 0.1654 | ||
| 274 | 0.0185 | ||
| 276 | 0.2712 | ||
| 278 | 0.0298 | ||
| 22 | REN54P11 | 222 | 0.1683 |
| 226 | 0.2670 | ||
| 228 | 0.1104 | ||
| 232 | 0.0903 | ||
| 234 | 0.0600 | ||
| 236 | 0.0407 | ||
| 238 | 0.2632 | ||
| 23 | REN64E19 | 139 | 0.1423 |
| 143 | 0.0130 | ||
| 145 | 0.0974 | ||
| 147 | 0.3107 | ||
| 149 | 0.0151 | ||
| 151 | 0.0038 | ||
| 153 | 0.3997 | ||
| 155 | 0.0176 | ||
| 157 | 0.0004 | ||
| 24 | VGL0760 | 12 | 0.0319 |
| 13 | 0.0609 | ||
| 14 | 0.1029 | ||
| 15 | 0.0097 | ||
| 18.2 | 0.0978 | ||
| 19.2 | 0.0025 | ||
| 20.2 | 0.0168 | ||
| 21.2 | 0.2330 | ||
| 22.2 | 0.1709 | ||
| 23.2 | 0.1612 | ||
| 24.2 | 0.0504 | ||
| 25.2 | 0.0290 | ||
| 26.2 | 0.0327 | ||
| 27.2 | 0.0004 | ||
| 25 | VGL0910 | 13 | 0.0038 |
| 14 | 0.0097 | ||
| 16.1 | 0.0919 | ||
| 17 | 0.0063 | ||
| 17.1 | 0.1012 | ||
| 18 | 0.0248 | ||
| 18.1 | 0.0609 | ||
| 19.1 | 0.1700 | ||
| 20.1 | 0.0407 | ||
| 20.3 | 0.0050 | ||
| 21.1 | 0.3149 | ||
| 22 | 0.0327 | ||
| 22.1 | 0.1062 | ||
| 23 | 0.0155 | ||
| 23.1 | 0.0164 | ||
| 26 | VGL1063 | 8 | 0.0235 |
| 12 | 0.0844 | ||
| 13 | 0.0399 | ||
| 14 | 0.1982 | ||
| 15 | 0.1029 | ||
| 16 | 0.0071 | ||
| 17 | 0.0877 | ||
| 18 | 0.0688 | ||
| 19 | 0.0437 | ||
| 20 | 0.0856 | ||
| 21 | 0.0894 | ||
| 22 | 0.1562 | ||
| 23 | 0.0126 | ||
| 27 | VGL1165 | 14 | 0.0008 |
| 15 | 0.0004 | ||
| 16 | 0.0059 | ||
| 17 | 0.0294 | ||
| 18 | 0.1108 | ||
| 19 | 0.0155 | ||
| 20 | 0.0290 | ||
| 21 | 0.1125 | ||
| 22 | 0.0004 | ||
| 23 | 0.0248 | ||
| 25 | 0.0038 | ||
| 26 | 0.3526 | ||
| 27 | 0.0705 | ||
| 28 | 0.1746 | ||
| 29 | 0.0080 | ||
| 30 | 0.0113 | ||
| 31 | 0.0231 | ||
| 32 | 0.0105 | ||
| 33 | 0.0151 | ||
| 34 | 0.0008 | ||
| 28 | VGL1828 | 15 | 0.0143 |
| 16 | 0.3073 | ||
| 17 | 0.0050 | ||
| 18 | 0.0214 | ||
| 19 | 0.0319 | ||
| 19.3 | 0.0029 | ||
| 20 | 0.1096 | ||
| 20.3 | 0.0004 | ||
| 21 | 0.3606 | ||
| 22 | 0.0638 | ||
| 23 | 0.0819 | ||
| 24 | 0.0008 | ||
| 29 | VGL2009 | 9 | 0.0021 |
| 10 | 0.0025 | ||
| 11 | 0.1457 | ||
| 12 | 0.0256 | ||
| 13 | 0.2145 | ||
| 14 | 0.2284 | ||
| 15 | 0.2939 | ||
| 16 | 0.0861 | ||
| 17 | 0.0013 | ||
| 30 | VGL2409 | 14 | 0.0118 |
| 15 | 0.0701 | ||
| 16 | 0.3279 | ||
| 17 | 0.3241 | ||
| 18 | 0.2120 | ||
| 19 | 0.0029 | ||
| 20 | 0.0420 | ||
| 21 | 0.0092 | ||
| 31 | VGL2918 | 12 | 0.0441 |
| 13 | 0.2821 | ||
| 14 | 0.1486 | ||
| 15 | 0.0680 | ||
| 16 | 0.0109 | ||
| 16.3 | 0.0055 | ||
| 17.3 | 0.0109 | ||
| 18.3 | 0.0668 | ||
| 19.3 | 0.0411 | ||
| 20.3 | 0.0306 | ||
| 21.3 | 0.0672 | ||
| 22.3 | 0.2158 | ||
| 23.3 | 0.0084 | ||
| 32 | VGL3008 | 10 | 0.0004 |
| 13 | 0.1276 | ||
| 14 | 0.0084 | ||
| 15 | 0.0709 | ||
| 16 | 0.0264 | ||
| 17 | 0.1100 | ||
| 18 | 0.1537 | ||
| 19 | 0.1767 | ||
| 20 | 0.2040 | ||
| 21 | 0.1041 | ||
| 22 | 0.0105 | ||
| 23 | 0.0071 | ||
| 33 | VGL3235 | 11 | 0.0109 |
| 12 | 0.0181 | ||
| 13 | 0.1264 | ||
| 14 | 0.2783 | ||
| 15 | 0.3438 | ||
| 16 | 0.0285 | ||
| 17 | 0.0395 | ||
| 18 | 0.0663 | ||
| 19 | 0.0835 | ||
| 20 | 0.0046 |
Standard genetic assessment based on 33 autosomal STR loci
| N | Na | Ne | Ho | He | F | ||
|---|---|---|---|---|---|---|---|
| Mean | 1191 | 9.515 | 4.475 | 0.730 | 0.747 | 0.023 | |
| SE | 0.535 | 0.268 | 0.016 | 0.016 | 0.004 |
Standard genetic assessment based on 7 STRs in the DLA region
| N | Na | Ne | Ho | He | F | ||
|---|---|---|---|---|---|---|---|
| Mean | 1191 | 8.857 | 3.837 | 0.669 | 0.690 | 0.032 | |
| SE | 0.999 | 0.600 | 0.047 | 0.047 | 0.005 |
Standard genetic assessment for individual STR loci
Havanese
| # | Locus | N | Na | Ne | Ho | He | F |
|---|---|---|---|---|---|---|---|
| 1 | AHT121 | 1191 | 12 | 3.119 | 0.673 | 0.679 | 0.009 |
| 2 | AHT137 | 1191 | 11 | 6.422 | 0.832 | 0.844 | 0.014 |
| 3 | AHTH130 | 1191 | 9 | 4.048 | 0.715 | 0.753 | 0.050 |
| 4 | AHTh171-A | 1191 | 10 | 4.841 | 0.787 | 0.793 | 0.008 |
| 5 | AHTh260 | 1191 | 9 | 4.840 | 0.769 | 0.793 | 0.031 |
| 6 | AHTk211 | 1191 | 5 | 3.149 | 0.652 | 0.682 | 0.045 |
| 7 | AHTk253 | 1191 | 5 | 2.117 | 0.485 | 0.528 | 0.080 |
| 8 | C22.279 | 1191 | 8 | 4.632 | 0.757 | 0.784 | 0.034 |
| 9 | FH2001 | 1191 | 9 | 2.630 | 0.625 | 0.620 | -0.008 |
| 10 | FH2054 | 1191 | 10 | 6.189 | 0.831 | 0.838 | 0.009 |
| 11 | FH2848 | 1191 | 9 | 4.409 | 0.752 | 0.773 | 0.027 |
| 12 | INRA21 | 1191 | 7 | 4.428 | 0.743 | 0.774 | 0.040 |
| 13 | INU005 | 1191 | 9 | 2.456 | 0.594 | 0.593 | -0.003 |
| 14 | INU030 | 1191 | 7 | 2.238 | 0.544 | 0.553 | 0.016 |
| 15 | INU055 | 1191 | 6 | 4.113 | 0.730 | 0.757 | 0.036 |
| 16 | LEI004 | 1191 | 9 | 2.105 | 0.533 | 0.525 | -0.016 |
| 17 | REN105L03 | 1191 | 9 | 3.455 | 0.683 | 0.711 | 0.038 |
| 18 | REN162C04 | 1191 | 6 | 3.920 | 0.730 | 0.745 | 0.019 |
| 19 | REN169D01 | 1191 | 8 | 3.907 | 0.733 | 0.744 | 0.015 |
| 20 | REN169O18 | 1191 | 7 | 5.183 | 0.793 | 0.807 | 0.017 |
| 21 | REN247M23 | 1191 | 7 | 3.548 | 0.703 | 0.718 | 0.021 |
| 22 | REN54P11 | 1191 | 7 | 5.141 | 0.772 | 0.805 | 0.041 |
| 23 | REN64E19 | 1191 | 9 | 3.488 | 0.712 | 0.713 | 0.002 |
| 24 | VGL0760 | 1191 | 14 | 7.185 | 0.851 | 0.861 | 0.012 |
| 25 | VGL0910 | 1191 | 15 | 6.033 | 0.814 | 0.834 | 0.025 |
| 26 | VGL1063 | 1191 | 13 | 8.820 | 0.877 | 0.887 | 0.011 |
| 27 | VGL1165 | 1191 | 20 | 5.307 | 0.762 | 0.812 | 0.062 |
| 28 | VGL1828 | 1191 | 12 | 4.016 | 0.766 | 0.751 | -0.020 |
| 29 | VGL2009 | 1191 | 9 | 4.677 | 0.759 | 0.786 | 0.035 |
| 30 | VGL2409 | 1191 | 8 | 3.782 | 0.713 | 0.736 | 0.031 |
| 31 | VGL2918 | 1191 | 13 | 5.997 | 0.827 | 0.833 | 0.007 |
| 32 | VGL3008 | 1191 | 12 | 7.059 | 0.848 | 0.858 | 0.012 |
| 33 | VGL3235 | 1191 | 10 | 4.427 | 0.732 | 0.774 | 0.054 |
Standard genetic assessment for 7 STRs in the DLA region
Havanese
| # | Locus | N | Na | Ne | Ho | He | F |
|---|---|---|---|---|---|---|---|
| 1 | DLA I-3CCA | 1191 | 12 | 5.036 | 0.782 | 0.801 | 0.025 |
| 2 | DLA I-4ACA | 1191 | 12 | 4.700 | 0.759 | 0.787 | 0.036 |
| 3 | DLA I-4BCT | 1191 | 8 | 2.039 | 0.500 | 0.510 | 0.020 |
| 4 | DLA1131 | 1191 | 11 | 6.663 | 0.833 | 0.850 | 0.020 |
| 5 | 5ACA | 1191 | 6 | 3.581 | 0.699 | 0.721 | 0.031 |
| 6 | 5ACT | 1191 | 5 | 2.130 | 0.499 | 0.531 | 0.060 |
| 7 | 5BCA | 1191 | 8 | 2.710 | 0.610 | 0.631 | 0.034 |